Nigel Farage has claimed his party will win double the number of seats predicted as UKIP makes significant gains across the country.
Early on Friday morning UKIP had already surpassed the 80 seats it had been expected to take, dealing a significant blow to the main parties.
UKIP took seats off both Labour and the Conservatives in their heartlands, which was particularly damaging to Ed Miliband, who has been accused by his own party of running a "unforgiveably unprofessional" campaign.
A ballot box is emptied at Trinity School in CroydonLabour lost its grip in the north and, in a serious blow, the party lost control of Thurrock, a key marginal for the General Election in 2015, to no overall control, losing two seats to UKIP.
Mr Farage said UKIP would now be "serious players" in the General Election and said the party was expecting to win double the 80 seats that had been predicted.
He said: "The UKIP fox is in the Westminster henhouse" and added: "The idea the UKIP vote just hurts the Tories is going to be blown away by this election."
However, Mr Farage, who has consistently warned the local and European elections would deliver a UKIP "earthquake", admitted that the party was unlikely to be successful in London.
Douglas Alexander, Labour's election co-ordinator, was forced to deny fighting a "lacklustre" campaign.
He told Sky News the only way to tackle the march of UKIP was by deploying an "army of activists" on the doorsteps and said that Labour had knocked on seven million doors.
Joey Essex may have supported Ed Miliband but Essex man voted UKIPMr Alexander said there was a "deep antipathy to how politics has been done" and UKIP had scooped up that vote but that "strengthening and growing the ground operation" was the only strategy for 2015.
UKIP made its greatest gains in Essex, where Margaret Thatcher once identified the "Essex Man", a man who moved out of London, once voted Labour but switched to the Tories.
UKIP took seats from Labour in Hartlepool, won 10 seats in Rotherham and polled more than a third of the vote in wards in big cities, such as Sunderland, Birmingham and Hull, where it previously had little or no presence.
According to the latest Sky News projection, the results so far would give a hung parliament at the 2015 General Election.
Mr Farage casts his vote in Cudham, KentSky's election analyst Professor Michael Thrasher said UKIP's success suggested the party would claim at least one seat in the House of Commons next year
Conservative Education Secretary Michael Gove said the results had not been as bad as expected. He categorically ruled out any chance of a pact with UKIP.
Most councils will not declare their results until later on Friday.
The Camerons and the Milibands cast their votesAs predicted, the Liberal Democrats suffered significant losses, particularly in Portsmouth where it lost control with UKIP gaining six seats.
However, it managed to hold on in Eastleigh, where UKIP had been expected to make gains, which was a significant victory for the party.
Business Secretary Vince Cable admitted it would be a bad night, adding: "We take a kicking for the things that government does that are unpopular."
:: Follow all the results as they come in on Twitter with @skyelections.
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